Sean,
In theory you can look at what WAS built, and WHY it was built, no?
The links land was actually used mostly because it was deemed unfit for other development purposes, no? Similarly, open inland sites were valued in many cases to avoid the cost of heavy clearing. Only later did tech allow golf to be built on wooded sites. IMHO, golf is adaptable with the proper technology, and agree that variation is the spice of life, and of golf courses. There is no right way or right site, although those "unsuited" ocean sites were certainly a god send to golf.
It would appear, as TD says, that golfers prefer some connection to nature, though. And, in abscence of water, trees appear to be the next best thing. IMHO, they also generally prefer some separation between holes, unless anyone can give a reason all those open courses were planted so heavily over the years.
It might have been for safety or other reason. However, people do enjoy being in "confined or defined" spaces. Hence the routing in dunes land mostly in the valleys. It so happens that the space of one hole makes for a nicely and humanly scaled defined space, also used so often by the Enlgish Estates - the open meadow with a defined end..
But, its not universal either - broad vistas can easily trump enclosed spaces, so if the vista is broad enough, its probably more satisfying to most than the defined space. Also, I have used the old landscape architecture trick of designing views "around the corner" or "peep hole" views that give a combo of defined space vs connection to the rest of the world.