Tim,
I would bet a bunch of money that the WS maintenance budget is very close to that of the courses your named above. Now, they may not spend as much time getting the greens fast, but there is A LOT of handwork there-greens are walk mowed, as are tees and approaches, bunkers are hand raked, they handwater, and there is A LOT of detail work. They carry a staff of 35. With greens fees of $340, plus $60 for a caddie, their conditions are as good as many private clubs.