If I may, I'd like to recommend a book on this subject. It's called "Biomimicry," by a scientist (I couldn't tell you what specific discipline, but somewhere in the life sciences) named Janine Benyus.
She argues that the built environment could be better designed if we absorbed everyday lessons (such as the strength and simplicity of a spider's web) from the natural world and applied them to our designs.
So, perhaps golf, and golf course design, is cannot be summed up in an equation, but surely we can all benefit from the sometimes subtle education of the natural environment.