Matt
We really need Mark Chaplin and Craig Disher to chime in. They know a lot more than me!
Here's what Rye's got, ex the 3s, and including the only par 5 (the 1st) as it in fact may play more like a 4 than the 437-yard third (which sometimes felt like more like a par 5 to me). Distances in yards:
Front
Back
Of course, on a links with no irrigation, these yardages don't tell the entire story. It is a shorter course in High Summer.
To Michael's point we don't play a course 6,000 yards at a time, we play it one hole at a time. Because the course has five par 3s, two short par 4s, and just one par 5, that leaves 10 holes to cover a lot of distance. (Throw in the 222-yard 17th and that's nearly one mile of golf to negotiate in the final four holes against a par of 15.)
Matt, I understand you are a long hitter and so maybe things are different for Padraig and you but for me these holes offered plenty of variety.
I can remember playing each of the holes listed above as though it were yesterday. I found 8, 10, and 11 the least interesting; 12, 15, 16 and possibly 18 I thought were fantastic. 4 is all-world! (8, 10, and 11 have the weakest relationship to the prominent feature of the course: a long dune.) 12, 15, and 16 form a rough triangle while 4 and 18 run in opposite directions, so: these four holes will play differently no matter the conditions.
In comparison to other top UK courses I've played, the par 4s hold up I think quite well. It's not as "great" a test of driving as, say, Woodhall Spa, but then Rye's a far more subtle type of test. It doesn't dictate the test like Woodhall Spa does; you really have to reason back from the greens -- the complexes are fantastic in a sublime way -- and even think ahead several holes to decide how to play the hole in question. For example, the short 9th: it's a good idea to think about your odds on 12 and 13 when deciding whether to play aggressively.
Another comparison: if I had to pick between Ailsa and Rye, I'd probably go 7 rounds Rye, 3 rounds Turnberry. Certainly no worse than 6-4. Put a screen up around each course's perimeter and it probably goes 8-2.
Does this help?
Mark